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[[File:Stromatolites in Sharkbay.jpg|thumb|Stromatolites in [[Shark Bay]] 鲨鱼湾中的石笋]]
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[[File:Stromatolites in Sharkbay.jpg|thumb|鲨鱼湾中的石笋]]
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The earliest life on Earth existed more than 3.5 Gya (billion years ago),<ref name="Origin1" /><ref name="Origin2" /><ref name="RavenJohnson2002" /> during the [[Eoarchean]] Era when sufficient crust had solidified following the molten Hadean Eon. The earliest physical evidence so far found consists of [[Micropaleontology#Microfossils|microfossils]] in the [[Nuvvuagittuq Greenstone Belt]] of Northern Quebec, in [[banded iron formation]] rocks at least 3.77 and possibly 4.28&nbsp;Gyo.<ref name="NAT-20170301" /><ref>{{cite web |url=http://www.cbc.ca/news/technology/oldest-record-life-earth-found-quebec-1.4004545 |title=Oldest traces of life on Earth found in Quebec, dating back roughly 3.8&nbsp;Gya |author=Mortillaro, Nicole |publisher=CBC News |date=1 March 2017 |accessdate=2 March 2017 |url-status=live |archiveurl=https://web.archive.org/web/20170301221842/http://www.cbc.ca/news/technology/oldest-record-life-earth-found-quebec-1.4004545 |archivedate=1 March 2017}}</ref> This finding suggested life developed very soon after oceans formed. The structure of the microbes was noted to be similar to bacteria found near [[hydrothermal vents]] in the modern era, and provided support for the hypothesis that abiogenesis began near hydrothermal vents.<ref name="4.3b oldest" /><ref name="NAT-20170301" />
      
地球上最早的生命存在于35亿年前,<ref name="Origin1" /><ref name="Origin2" /><ref name="RavenJohnson2002" />在始太古代时期,在熔化的冥古宙之后,充足的地壳已经凝固。迄今为止发现的最早的物理证据包括魁北克北部努夫雅集图克绿岩带中的微生物化石,位于至少37.7亿岁,也可能42.8亿岁的带状铁层岩石中。<ref name="NAT-20170301" /><ref>{{cite web |url=http://www.cbc.ca/news/technology/oldest-record-life-earth-found-quebec-1.4004545 |title=Oldest traces of life on Earth found in Quebec, dating back roughly 3.8&nbsp;Gya |author=Mortillaro, Nicole |publisher=CBC News |date=1 March 2017 |accessdate=2 March 2017 |url-status=live |archiveurl=https://web.archive.org/web/20170301221842/http://www.cbc.ca/news/technology/oldest-record-life-earth-found-quebec-1.4004545 |archivedate=1 March 2017}}</ref>这个发现表明生命在海洋形成后不久便出现了。据悉,这种微生物的结构与现代热液喷口附近发现的细菌相似,为无生源论始于热液喷口附近的假说提供了支持。
 
地球上最早的生命存在于35亿年前,<ref name="Origin1" /><ref name="Origin2" /><ref name="RavenJohnson2002" />在始太古代时期,在熔化的冥古宙之后,充足的地壳已经凝固。迄今为止发现的最早的物理证据包括魁北克北部努夫雅集图克绿岩带中的微生物化石,位于至少37.7亿岁,也可能42.8亿岁的带状铁层岩石中。<ref name="NAT-20170301" /><ref>{{cite web |url=http://www.cbc.ca/news/technology/oldest-record-life-earth-found-quebec-1.4004545 |title=Oldest traces of life on Earth found in Quebec, dating back roughly 3.8&nbsp;Gya |author=Mortillaro, Nicole |publisher=CBC News |date=1 March 2017 |accessdate=2 March 2017 |url-status=live |archiveurl=https://web.archive.org/web/20170301221842/http://www.cbc.ca/news/technology/oldest-record-life-earth-found-quebec-1.4004545 |archivedate=1 March 2017}}</ref>这个发现表明生命在海洋形成后不久便出现了。据悉,这种微生物的结构与现代热液喷口附近发现的细菌相似,为无生源论始于热液喷口附近的假说提供了支持。
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Biogenic [[graphite]] has been found in 3.7&nbsp;Gyo metasedimentary rocks from southwestern [[Greenland]]<ref name="NG-20131208">{{cite journal |last1=Ohtomo |first1=Yoko |last2=Kakegawa |first2=Takeshi |last3=Ishida |first3=Akizumi |last4=Nagase |first4=Toshiro |last5=Rosing |first5=Minik T. |display-authors=3 |date=January 2014 |title=Evidence for biogenic graphite in early Archaean Isua metasedimentary rocks |journal=[[Nature Geoscience]] |volume=7 |issue=1 |pages=25–28 |bibcode=2014NatGe...7...25O |doi=10.1038/ngeo2025 }}</ref> and [[microbial mat]] fossils found in 3.48&nbsp;Gyo sandstone from [[Western Australia]].<ref name="AP-20131113">{{cite news |last=Borenstein |first=Seth |date=13 November 2013 |title=Oldest fossil found: Meet your microbial mom |url=http://apnews.excite.com/article/20131113/DAA1VSC01.html |work=[[Excite]] |location=Yonkers, NY |publisher=[[Mindspark Interactive Network]] |agency=[[Associated Press]] |accessdate=2015-06-02 |url-status=live |archiveurl=https://web.archive.org/web/20150629230719/http://apnews.excite.com/article/20131113/DAA1VSC01.html |archivedate=29 June 2015}}</ref><ref name="AST-20131108">{{cite journal |last1=Noffke |first1=Nora |last2=Christian |first2=Daniel |last3=Wacey |first3=David |last4=Hazen |first4=Robert M. |authorlink4=Robert Hazen |date=16 November 2013 |title=Microbially Induced Sedimentary Structures Recording an Ancient Ecosystem in the ''ca.'' 3.48 Gyo Dresser Formation, Pilbara, Western Australia |journal=[[Astrobiology (journal)|Astrobiology]] |volume=13 |issue=12 |pages=1103–1124 |bibcode=2013AsBio..13.1103N |doi=10.1089/ast.2013.1030 |pmc=3870916 |pmid=24205812}}</ref> Evidence of early life in rocks from [[Akilia]] Island, near the [[Isua Greenstone Belt|Isua supracrustal belt]] in southwestern Greenland, dating to 3.7&nbsp;Gya have shown biogenic [[carbon isotope]]s.<ref name="NYT-20160831">{{cite news |last=Wade |first=Nicholas |title=World's Oldest Fossils Found in Greenland |url=https://www.nytimes.com/2016/09/01/science/oldest-fossils-on-earth.html |date=31 August 2016 |work=[[The New York Times]] |accessdate=31 August 2016 |url-status=live |archiveurl=https://web.archive.org/web/20160831185959/http://www.nytimes.com/2016/09/01/science/oldest-fossils-on-earth.html |archivedate=31 August 2016}}</ref><ref>{{harvnb|Davies|1999}}</ref> In other parts of the Isua supracrustal belt, graphite inclusions trapped within [[garnet]] crystals are connected to the other elements of life: oxygen, nitrogen, and possibly phosphorus in the form of [[phosphate]], providing further evidence for life 3.7&nbsp;Gya.<ref>{{Cite journal |last1=Hassenkam|first1=T. |last2=Andersson |first2=M.P. |last3=Dalby|first3=K.N. |last4=Mackenzie |first4=D.M.A.|last5=Rosing |first5=M.T. |title=Elements of Eoarchean life trapped in mineral inclusions |journal=Nature |doi=10.1038/nature23261 |pmid=28738409 |volume=548|issue=7665|pages=78–81 |year=2017 |bibcode=2017Natur.548...78H|s2cid=205257931 }}</ref> At Strelley Pool, in the [[Pilbara]] region of Western Australia, compelling evidence of early life was found in [[pyrite]]-bearing sandstone in a fossilized beach, that showed rounded tubular cells that [[Redox|oxidized]] sulfur by [[photosynthesis]] in the absence of oxygen.<ref name="TG-20131113-JP">{{cite news |last=Pearlman |first=Jonathan |date=13 November 2013 |title=Oldest signs of life on Earth found |url=https://www.telegraph.co.uk/news/science/science-news/10445788/Oldest-signs-of-life-on-Earth-found.html |newspaper=[[The Daily Telegraph]] |location=London |accessdate=2014-12-15 |url-status=live |archiveurl=https://web.archive.org/web/20141216062531/http://www.telegraph.co.uk/news/science/science-news/10445788/Oldest-signs-of-life-on-Earth-found.html |archivedate=16 December 2014}}</ref><ref>{{cite journal |last=O'Donoghue |first=James |date=21 August 2011 |url=https://www.newscientist.com/article/dn20813-oldest-reliable-fossils-show-early-life-was-a-beach.html |title=Oldest reliable fossils show early life was a beach |journal=[[New Scientist]] |url-status=live |archiveurl=https://web.archive.org/web/20150630201918/http://www.newscientist.com/article/dn20813-oldest-reliable-fossils-show-early-life-was-a-beach.html |archivedate=30 June 2015|doi=10.1016/S0262-4079(11)62064-2 |volume=211 |page=13 }}</ref><ref>{{cite journal |last1=Wacey |first1=David |last2=Kilburn |first2=Matt R. |last3=Saunders |first3=Martin |last4=Cliff |first4=John |last5=Brasier |first5=Martin D. |authorlink5=Martin Brasier |display-authors=3 |date=October 2011 |title=Microfossils of sulphur-metabolizing cells in 3.4-billion-year-old rocks of Western Australia |journal=Nature Geoscience |volume=4 |issue=10 |pages=698–702 |bibcode=2011NatGe...4..698W |doi=10.1038/ngeo1238}}</ref> Further research on [[zircon]]s from Western Australia in 2015 suggested that life likely existed on Earth at least 4.1 Gya.<ref name="AP-20151019">{{cite news |last=Borenstein |first=Seth |title=Hints of life on what was thought to be desolate early Earth |url=https://apnews.com/e6be2537b4cd46ffb9c0585bae2b2e51 |date=19 October 2015 |work=AP News |publisher=[[Associated Press]] |accessdate=9 October 2018}}</ref><ref name="PNAS-20151014-pdf">{{cite journal |last1=Bell |first1=Elizabeth A. |last2=Boehnike |first2=Patrick |last3=Harrison |first3=T. Mark |last4=Mao |first4=Wendy L. |display-authors=3 |date=19 October 2015 |title=Potentially biogenic carbon preserved in a 4.1 billion-year-old zircon|journal=Proc. Natl. Acad. Sci. U.S.A. |doi=10.1073/pnas.1517557112|pages=14518–14521 |pmid=26483481 |pmc=4664351 |volume=112 |issue=47 |bibcode=2015PNAS..11214518B}} Early edition, published online before print.</ref><ref name="UCLA-20151019">{{cite web |last1=Wolpert |first1=Stuart |title=Life on Earth likely started at least 4.1 billion years ago – much earlier than scientists had thought |url=http://newsroom.ucla.edu/releases/life-on-earth-likely-started-at-least-4-1-billion-years-ago-much-earlier-than-scientists-had-thought |date=19 October 2015 |publisher=[[ULCA]] |accessdate=20 October 2015 |url-status=live |archiveurl=https://web.archive.org/web/20151020164038/http://newsroom.ucla.edu/releases/life-on-earth-likely-started-at-least-4-1-billion-years-ago-much-earlier-than-scientists-had-thought |archivedate=20 October 2015}}</ref>
      
人们在格陵兰岛西南部37亿岁的变质沉积岩中发现了生物来源的石墨<ref name="NG-20131208">{{cite journal |last1=Ohtomo |first1=Yoko |last2=Kakegawa |first2=Takeshi |last3=Ishida |first3=Akizumi |last4=Nagase |first4=Toshiro |last5=Rosing |first5=Minik T. |display-authors=3 |date=January 2014 |title=Evidence for biogenic graphite in early Archaean Isua metasedimentary rocks |journal=[[Nature Geoscience]] |volume=7 |issue=1 |pages=25–28 |bibcode=2014NatGe...7...25O |doi=10.1038/ngeo2025 }}</ref> ,在西澳大利亚距今34.8亿年前的砂岩中发现了微生物垫层化石<ref name="AP-20131113">{{cite news |last=Borenstein |first=Seth |date=13 November 2013 |title=Oldest fossil found: Meet your microbial mom |url=http://apnews.excite.com/article/20131113/DAA1VSC01.html |work=[[Excite]] |location=Yonkers, NY |publisher=[[Mindspark Interactive Network]] |agency=[[Associated Press]] |accessdate=2015-06-02 |url-status=live |archiveurl=https://web.archive.org/web/20150629230719/http://apnews.excite.com/article/20131113/DAA1VSC01.html |archivedate=29 June 2015}}</ref><ref name="AST-20131108">{{cite journal |last1=Noffke |first1=Nora |last2=Christian |first2=Daniel |last3=Wacey |first3=David |last4=Hazen |first4=Robert M. |authorlink4=Robert Hazen |date=16 November 2013 |title=Microbially Induced Sedimentary Structures Recording an Ancient Ecosystem in the ''ca.'' 3.48 Gyo Dresser Formation, Pilbara, Western Australia |journal=[[Astrobiology (journal)|Astrobiology]] |volume=13 |issue=12 |pages=1103–1124 |bibcode=2013AsBio..13.1103N |doi=10.1089/ast.2013.1030 |pmc=3870916 |pmid=24205812}}</ref>。在格陵兰岛西南部伊苏亚上地壳带附近的阿基利亚岛的岩石中发现了早期生命的证据,这些可追溯到37亿年前的证据中发现了生源碳同位素<ref name="NYT-20160831">{{cite news |last=Wade |first=Nicholas |title=World's Oldest Fossils Found in Greenland |url=https://www.nytimes.com/2016/09/01/science/oldest-fossils-on-earth.html |date=31 August 2016 |work=[[The New York Times]] |accessdate=31 August 2016 |url-status=live |archiveurl=https://web.archive.org/web/20160831185959/http://www.nytimes.com/2016/09/01/science/oldest-fossils-on-earth.html |archivedate=31 August 2016}}</ref><ref>{{harvnb|Davies|1999}}</ref> 。在伊苏亚上地壳带的其他地方,被困在石榴石晶体内的石墨包裹体与生命的其他元素相连:氧、氮和可能以磷酸盐形式存在的磷,为37亿年前的生命提供了进一步的证据<ref>{{Cite journal |last1=Hassenkam|first1=T. |last2=Andersson |first2=M.P. |last3=Dalby|first3=K.N. |last4=Mackenzie |first4=D.M.A.|last5=Rosing |first5=M.T. |title=Elements of Eoarchean life trapped in mineral inclusions |journal=Nature |doi=10.1038/nature23261 |pmid=28738409 |volume=548|issue=7665|pages=78–81 |year=2017 |bibcode=2017Natur.548...78H|s2cid=205257931 }}</ref> 。在西澳大利亚皮尔巴拉地区的斯特雷利池,在一个化石滩的含黄铁矿砂岩中发现了早期生命的令人信服的证据,它显示了圆形的管状细胞,在没有氧气的情况下通过光合作用氧化硫。2015年对西澳大利亚的锆石的进一步研究表明,生命很可能在至少41亿年前就存在于地球上。<ref name="AP-20151019">{{cite news |last=Borenstein |first=Seth |title=Hints of life on what was thought to be desolate early Earth |url=https://apnews.com/e6be2537b4cd46ffb9c0585bae2b2e51 |date=19 October 2015 |work=AP News |publisher=[[Associated Press]] |accessdate=9 October 2018}}</ref><ref name="PNAS-20151014-pdf">{{cite journal |last1=Bell |first1=Elizabeth A. |last2=Boehnike |first2=Patrick |last3=Harrison |first3=T. Mark |last4=Mao |first4=Wendy L. |display-authors=3 |date=19 October 2015 |title=Potentially biogenic carbon preserved in a 4.1 billion-year-old zircon|journal=Proc. Natl. Acad. Sci. U.S.A. |doi=10.1073/pnas.1517557112|pages=14518–14521 |pmid=26483481 |pmc=4664351 |volume=112 |issue=47 |bibcode=2015PNAS..11214518B}} Early edition, published online before print.</ref><ref name="UCLA-20151019">{{cite web |last1=Wolpert |first1=Stuart |title=Life on Earth likely started at least 4.1 billion years ago – much earlier than scientists had thought |url=http://newsroom.ucla.edu/releases/life-on-earth-likely-started-at-least-4-1-billion-years-ago-much-earlier-than-scientists-had-thought |date=19 October 2015 |publisher=[[ULCA]] |accessdate=20 October 2015 |url-status=live |archiveurl=https://web.archive.org/web/20151020164038/http://newsroom.ucla.edu/releases/life-on-earth-likely-started-at-least-4-1-billion-years-ago-much-earlier-than-scientists-had-thought |archivedate=20 October 2015}}</ref>
 
人们在格陵兰岛西南部37亿岁的变质沉积岩中发现了生物来源的石墨<ref name="NG-20131208">{{cite journal |last1=Ohtomo |first1=Yoko |last2=Kakegawa |first2=Takeshi |last3=Ishida |first3=Akizumi |last4=Nagase |first4=Toshiro |last5=Rosing |first5=Minik T. |display-authors=3 |date=January 2014 |title=Evidence for biogenic graphite in early Archaean Isua metasedimentary rocks |journal=[[Nature Geoscience]] |volume=7 |issue=1 |pages=25–28 |bibcode=2014NatGe...7...25O |doi=10.1038/ngeo2025 }}</ref> ,在西澳大利亚距今34.8亿年前的砂岩中发现了微生物垫层化石<ref name="AP-20131113">{{cite news |last=Borenstein |first=Seth |date=13 November 2013 |title=Oldest fossil found: Meet your microbial mom |url=http://apnews.excite.com/article/20131113/DAA1VSC01.html |work=[[Excite]] |location=Yonkers, NY |publisher=[[Mindspark Interactive Network]] |agency=[[Associated Press]] |accessdate=2015-06-02 |url-status=live |archiveurl=https://web.archive.org/web/20150629230719/http://apnews.excite.com/article/20131113/DAA1VSC01.html |archivedate=29 June 2015}}</ref><ref name="AST-20131108">{{cite journal |last1=Noffke |first1=Nora |last2=Christian |first2=Daniel |last3=Wacey |first3=David |last4=Hazen |first4=Robert M. |authorlink4=Robert Hazen |date=16 November 2013 |title=Microbially Induced Sedimentary Structures Recording an Ancient Ecosystem in the ''ca.'' 3.48 Gyo Dresser Formation, Pilbara, Western Australia |journal=[[Astrobiology (journal)|Astrobiology]] |volume=13 |issue=12 |pages=1103–1124 |bibcode=2013AsBio..13.1103N |doi=10.1089/ast.2013.1030 |pmc=3870916 |pmid=24205812}}</ref>。在格陵兰岛西南部伊苏亚上地壳带附近的阿基利亚岛的岩石中发现了早期生命的证据,这些可追溯到37亿年前的证据中发现了生源碳同位素<ref name="NYT-20160831">{{cite news |last=Wade |first=Nicholas |title=World's Oldest Fossils Found in Greenland |url=https://www.nytimes.com/2016/09/01/science/oldest-fossils-on-earth.html |date=31 August 2016 |work=[[The New York Times]] |accessdate=31 August 2016 |url-status=live |archiveurl=https://web.archive.org/web/20160831185959/http://www.nytimes.com/2016/09/01/science/oldest-fossils-on-earth.html |archivedate=31 August 2016}}</ref><ref>{{harvnb|Davies|1999}}</ref> 。在伊苏亚上地壳带的其他地方,被困在石榴石晶体内的石墨包裹体与生命的其他元素相连:氧、氮和可能以磷酸盐形式存在的磷,为37亿年前的生命提供了进一步的证据<ref>{{Cite journal |last1=Hassenkam|first1=T. |last2=Andersson |first2=M.P. |last3=Dalby|first3=K.N. |last4=Mackenzie |first4=D.M.A.|last5=Rosing |first5=M.T. |title=Elements of Eoarchean life trapped in mineral inclusions |journal=Nature |doi=10.1038/nature23261 |pmid=28738409 |volume=548|issue=7665|pages=78–81 |year=2017 |bibcode=2017Natur.548...78H|s2cid=205257931 }}</ref> 。在西澳大利亚皮尔巴拉地区的斯特雷利池,在一个化石滩的含黄铁矿砂岩中发现了早期生命的令人信服的证据,它显示了圆形的管状细胞,在没有氧气的情况下通过光合作用氧化硫。2015年对西澳大利亚的锆石的进一步研究表明,生命很可能在至少41亿年前就存在于地球上。<ref name="AP-20151019">{{cite news |last=Borenstein |first=Seth |title=Hints of life on what was thought to be desolate early Earth |url=https://apnews.com/e6be2537b4cd46ffb9c0585bae2b2e51 |date=19 October 2015 |work=AP News |publisher=[[Associated Press]] |accessdate=9 October 2018}}</ref><ref name="PNAS-20151014-pdf">{{cite journal |last1=Bell |first1=Elizabeth A. |last2=Boehnike |first2=Patrick |last3=Harrison |first3=T. Mark |last4=Mao |first4=Wendy L. |display-authors=3 |date=19 October 2015 |title=Potentially biogenic carbon preserved in a 4.1 billion-year-old zircon|journal=Proc. Natl. Acad. Sci. U.S.A. |doi=10.1073/pnas.1517557112|pages=14518–14521 |pmid=26483481 |pmc=4664351 |volume=112 |issue=47 |bibcode=2015PNAS..11214518B}} Early edition, published online before print.</ref><ref name="UCLA-20151019">{{cite web |last1=Wolpert |first1=Stuart |title=Life on Earth likely started at least 4.1 billion years ago – much earlier than scientists had thought |url=http://newsroom.ucla.edu/releases/life-on-earth-likely-started-at-least-4-1-billion-years-ago-much-earlier-than-scientists-had-thought |date=19 October 2015 |publisher=[[ULCA]] |accessdate=20 October 2015 |url-status=live |archiveurl=https://web.archive.org/web/20151020164038/http://newsroom.ucla.edu/releases/life-on-earth-likely-started-at-least-4-1-billion-years-ago-much-earlier-than-scientists-had-thought |archivedate=20 October 2015}}</ref>
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== 20世纪60年代以前的概念演变史:生物学 Conceptual history until the 1960s: biology ==
 
== 20世纪60年代以前的概念演变史:生物学 Conceptual history until the 1960s: biology ==
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