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道金斯一直对进化过程中的非适应性过程(如古尔德和路翁丁所描述的 Spandrels)以及“高于”基因水平的选择持怀疑态度<ref name="gould-lewontin">{{cite journal |last=Gould |first=Stephen Jay|author2=Lewontin, Richard C.  |year=1979 |title=The Spandrels of San Marco and the Panglossian Paradigm: A Critique of the Adaptationist Programme |journal=Proceedings of the Royal Society of London |volume=205 |issue=1161 |series=B |pages=581–98 |doi=10.1098/rspb.1979.0086 |pmid=42062 |bibcode=1979RSPSB.205..581G}}</ref> and about selection at levels "above" that of the gene.<ref name=Extended_Phenotype>{{cite book |last=Dawkins |first=Richard |title=The extended phenotype: the long reach of the gene |url=https://archive.org/details/extendedphenotyp0000dawk |url-access=registration |year=1999 |publisher=Oxford University Press |isbn=978-0192880512 |edition=Revised with new afterword and further reading}}</ref>。他特别怀疑群体选择作为理解利他主义的基础的实际可能性或重要性。这种行为起初似乎是一种进化悖论，因为帮助他人会耗费宝贵的资源，并降低自己的适应能力。以前，许多人把这解释为群体选择的一个方面：个体做的是对种群或整个物种的生存最有利的事情。英国进化生物学家W.D.汉密尔顿在他的整体适应度理论中使用基因频率分析来说明，如果这种利他主义的行为者和接受者(包括近亲)之间有足够的遗传相似性，那么遗传的利他主义特征是如何进化的<ref>{{cite journal |last=Hamilton |first=W.D.  |title=The genetical evolution of social behaviour I and II |journal=Journal of Theoretical Biology |volume=7 |issue=1 |pages=1–16, 17–52 |year=1964 |doi=10.1016/0022-5193(64)90038-4 |pmid=5875341}}</ref>。汉密尔顿的整体适应度已经成功地应用于包括人类在内的广泛的生物体中。类似地，罗伯特·泰弗士从以基因为中心的模型的角度思考，发展了互利主义理论，即一个有机体在对未来互换的预期中为另一个有机体提供利益<ref>{{cite journal |last=Trivers |first=Robert |title=The evolution of reciprocal altruism |journal=Quarterly Review of Biology |volume=46 |issue=1 |pages=35–57 |year=1971 |doi=10.1086/406755 |url=https://semanticscholar.org/paper/4e671994e5b0c7aefbecd050e95fdb45272d7e12 |access-date=16 December 2019 |archive-date=21 November 2020 |archive-url=https://web.archive.org/web/20201121060816/https://www.semanticscholar.org/paper/The-Evolution-of-Reciprocal-Altruism-Trivers/4e671994e5b0c7aefbecd050e95fdb45272d7e12 |url-status=live }}</ref>。道金斯在《自私的基因》中推广了这些观点，并在自己的作品中加以发展<ref name="dawkins79">{{cite journal |last=Dawkins |first=Richard |title=Twelve Misunderstandings of Kin Selection |journal=Zeitschrift für Tierpsychologie |volume=51 |pages=184–200 |year=1979 |issue=2 |url=http://www.simonyi.ox.ac.uk/dawkins/writings/Twelve%20Misunderstandings%20of%20Kin%20Selection.pdf |archive-url=https://web.archive.org/web/20080529180009/http://www.simonyi.ox.ac.uk/dawkins/writings/Twelve%20Misunderstandings%20of%20Kin%20Selection.pdf |archive-date=29 May 2008 |doi=10.1111/j.1439-0310.1979.tb00682.x}}</ref>。2012年6月，道金斯对生物学家 e. o. 威尔逊2012年出版的《地球的社会征服》一书提出严厉批评，认为该书误解了汉密尔顿的亲缘选择理论<ref>{{cite news |last=Thorpe |first=Vanessa |title=Richard Dawkins in furious row with EO Wilson over theory of evolution. Book review sparks war of words between grand old man of biology and Oxford's most high-profile Darwinist |url=https://www.theguardian.com/science/2012/jun/24/battle-of-the-professors |access-date=3 October 2012 |newspaper=The Guardian |date=24 June 2012 |location=London |archive-date=6 May 2019 |archive-url=https://web.archive.org/web/20190506014702/https://www.theguardian.com/science/2012/jun/24/battle-of-the-professors |url-status=live }}</ref><ref>{{cite news |last1=Dawkins |first1=Richard |title=The Descent of Edward Wilson |url=http://www.prospectmagazine.co.uk/magazine/edward-wilson-social-conquest-earth-evolutionary-errors-origin-species |access-date=24 October 2015 |work=Prospect |date=24 May 2012 |archive-date=24 September 2015 |archive-url=https://web.archive.org/web/20150924105332/http://www.prospectmagazine.co.uk/magazine/edward-wilson-social-conquest-earth-evolutionary-errors-origin-species |url-status=live }}</ref>。道金斯还强烈批评独立科学家詹姆斯·洛夫洛克的盖亚假说。<ref>{{cite book |title=The molecular biology of Gaia |url=https://archive.org/details/molecularbiology0000will |url-access=registration |year=1996 |publisher=Columbia University Press |isbn=978-0-231-10512-5 |page=[https://archive.org/details/molecularbiology0000will/page/178 178] |first1=George Ronald |last1=Williams}} [https://archive.org/details/molecularbiology0000will/page/178 Extract of page 178]</ref><ref>{{cite book |title=Scientists debate gaia: the next century |url=https://books.google.com/books?id=TOi1Cyj9h1UC |year=2004 |publisher=MIT Press |isbn=978-0-262-19498-3 |page=72 |first1=Stephen Henry |last1=Schneider |access-date=27 January 2016 |archive-date=29 July 2016 |archive-url=https://web.archive.org/web/20160729013112/https://books.google.com/books?id=TOi1Cyj9h1UC |url-status=live }{{Webarchive|url=https://web.archive.org/web/20150319005453/http://books.google.com/books?id=TOi1Cyj9h1UC&pg=PA72 |date=19 March 2015 }}</ref><ref>{{cite book |title=Unweaving the Rainbow: Science, Delusion and the Appetite for Wonder |journal=Unweaving the Rainbow : Science |url=https://books.google.com/books?id=ZudTchiioUoC |year=2000 |publisher=Houghton Mifflin Harcourt |isbn=978-0-618-05673-6 |page=223 |first1=Richard |last1=Dawkins |bibcode=1998ursd.book.....D |access-date=27 January 2016 |archive-date=21 September 2014 |archive-url=https://web.archive.org/web/20140921122549/http://books.google.com/books?id=ZudTchiioUoC |url-status=live }} [https://books.google.com/books?id=ZudTchiioUoC&pg=PA223 Extract of p. 223] {{Webarchive|url=https://web.archive.org/web/20150319064040/http://books.google.com/books?id=ZudTchiioUoC&pg=PA223 |date=19 March 2015 }}</ref>

道金斯一直对进化过程中的非适应性过程(如古尔德和路翁丁所描述的 Spandrels)以及“高于”基因水平的选择持怀疑态度<ref name="gould-lewontin">{{cite journal |last=Gould |first=Stephen Jay|author2=Lewontin, Richard C.  |year=1979 |title=The Spandrels of San Marco and the Panglossian Paradigm: A Critique of the Adaptationist Programme |journal=Proceedings of the Royal Society of London |volume=205 |issue=1161 |series=B |pages=581–98 |doi=10.1098/rspb.1979.0086 |pmid=42062 |bibcode=1979RSPSB.205..581G}}</ref> and about selection at levels "above" that of the gene.<ref name=Extended_Phenotype>{{cite book |last=Dawkins |first=Richard |title=The extended phenotype: the long reach of the gene |url=https://archive.org/details/extendedphenotyp0000dawk |url-access=registration |year=1999 |publisher=Oxford University Press |isbn=978-0192880512 |edition=Revised with new afterword and further reading}}</ref>。他特别怀疑群体选择作为理解利他主义的基础的实际可能性或重要性。这种行为起初似乎是一种进化悖论，因为帮助他人会耗费宝贵的资源，并降低自己的适应能力。以前，许多人把这解释为群体选择的一个方面：个体做的是对种群或整个物种的生存最有利的事情。英国进化生物学家W.D.汉密尔顿在他的整体适应度理论中使用基因频率分析来说明，如果这种利他主义的行为者和接受者(包括近亲)之间有足够的遗传相似性，那么遗传的利他主义特征是如何进化的<ref>{{cite journal |last=Hamilton |first=W.D.  |title=The genetical evolution of social behaviour I and II |journal=Journal of Theoretical Biology |volume=7 |issue=1 |pages=1–16, 17–52 |year=1964 |doi=10.1016/0022-5193(64)90038-4 |pmid=5875341}}</ref>。汉密尔顿的整体适应度已经成功地应用于包括人类在内的广泛的生物体中。类似地，罗伯特·泰弗士从以基因为中心的模型的角度思考，发展了互利主义理论，即一个有机体在对未来互换的预期中为另一个有机体提供利益<ref>{{cite journal |last=Trivers |first=Robert |title=The evolution of reciprocal altruism |journal=Quarterly Review of Biology |volume=46 |issue=1 |pages=35–57 |year=1971 |doi=10.1086/406755 |url=https://semanticscholar.org/paper/4e671994e5b0c7aefbecd050e95fdb45272d7e12 |access-date=16 December 2019 |archive-date=21 November 2020 |archive-url=https://web.archive.org/web/20201121060816/https://www.semanticscholar.org/paper/The-Evolution-of-Reciprocal-Altruism-Trivers/4e671994e5b0c7aefbecd050e95fdb45272d7e12 |url-status=live }}</ref>。道金斯在《自私的基因》中推广了这些观点，并在自己的作品中加以发展<ref name="dawkins79">{{cite journal |last=Dawkins |first=Richard |title=Twelve Misunderstandings of Kin Selection |journal=Zeitschrift für Tierpsychologie |volume=51 |pages=184–200 |year=1979 |issue=2 |url=http://www.simonyi.ox.ac.uk/dawkins/writings/Twelve%20Misunderstandings%20of%20Kin%20Selection.pdf |archive-url=https://web.archive.org/web/20080529180009/http://www.simonyi.ox.ac.uk/dawkins/writings/Twelve%20Misunderstandings%20of%20Kin%20Selection.pdf |archive-date=29 May 2008 |doi=10.1111/j.1439-0310.1979.tb00682.x}}</ref>。2012年6月，道金斯对生物学家 e. o. 威尔逊2012年出版的《地球的社会征服》一书提出严厉批评，认为该书误解了汉密尔顿的亲缘选择理论<ref>{{cite news |last=Thorpe |first=Vanessa |title=Richard Dawkins in furious row with EO Wilson over theory of evolution. Book review sparks war of words between grand old man of biology and Oxford's most high-profile Darwinist |url=https://www.theguardian.com/science/2012/jun/24/battle-of-the-professors |access-date=3 October 2012 |newspaper=The Guardian |date=24 June 2012 |location=London |archive-date=6 May 2019 |archive-url=https://web.archive.org/web/20190506014702/https://www.theguardian.com/science/2012/jun/24/battle-of-the-professors |url-status=live }}</ref><ref>{{cite news |last1=Dawkins |first1=Richard |title=The Descent of Edward Wilson |url=http://www.prospectmagazine.co.uk/magazine/edward-wilson-social-conquest-earth-evolutionary-errors-origin-species |access-date=24 October 2015 |work=Prospect |date=24 May 2012 |archive-date=24 September 2015 |archive-url=https://web.archive.org/web/20150924105332/http://www.prospectmagazine.co.uk/magazine/edward-wilson-social-conquest-earth-evolutionary-errors-origin-species |url-status=live }}</ref>。道金斯还强烈批评独立科学家詹姆斯·洛夫洛克的盖亚假说。<ref>{{cite book |title=The molecular biology of Gaia |url=https://archive.org/details/molecularbiology0000will |url-access=registration |year=1996 |publisher=Columbia University Press |isbn=978-0-231-10512-5 |page=[https://archive.org/details/molecularbiology0000will/page/178 178] |first1=George Ronald |last1=Williams}} [https://archive.org/details/molecularbiology0000will/page/178 Extract of page 178]</ref><ref>{{cite book |title=Scientists debate gaia: the next century |url=https://books.google.com/books?id=TOi1Cyj9h1UC |year=2004 |publisher=MIT Press |isbn=978-0-262-19498-3 |page=72 |first1=Stephen Henry |last1=Schneider |access-date=27 January 2016 |archive-date=29 July 2016 |archive-url=https://web.archive.org/web/20160729013112/https://books.google.com/books?id=TOi1Cyj9h1UC |url-status=live }} [https://books.google.com/books?id=TOi1Cyj9h1UC&pg=PA72 Extract of p. 72] {{Webarchive|url=https://web.archive.org/web/20150319005453/http://books.google.com/books?id=TOi1Cyj9h1UC&pg=PA72 |date=19 March 2015 }}</ref><ref>{{cite book |title=Unweaving the Rainbow: Science, Delusion and the Appetite for Wonder |journal=Unweaving the Rainbow : Science |url=https://books.google.com/books?id=ZudTchiioUoC |year=2000 |publisher=Houghton Mifflin Harcourt |isbn=978-0-618-05673-6 |page=223 |first1=Richard |last1=Dawkins |bibcode=1998ursd.book.....D |access-date=27 January 2016 |archive-date=21 September 2014 |archive-url=https://web.archive.org/web/20140921122549/http://books.google.com/books?id=ZudTchiioUoC |url-status=live }} [https://books.google.com/books?id=ZudTchiioUoC&pg=PA223 Extract of p. 223] {{Webarchive|url=https://web.archive.org/web/20150319064040/http://books.google.com/books?id=ZudTchiioUoC&pg=PA223 |date=19 March 2015 }}</ref>